Preheat oven to 350°F. Line an 8x8-inch square baking pan with parchment paper and spray with nonstick spray. Set aside.
In a large bowl, using a hand or stand mixer, beat together the cake mix, butter, egg, vanilla extract, lemon juice, and lemon zest until fully combined, about 30 seconds.
15.25 ounces lemon cake mix, ½ cup unsalted butter, 1 large egg, 1 teaspoon pure vanilla extract, 2 tablespoons fresh lemon juice, 1 tablespoon lemon zest
In a separate medium bowl, beat the cream cheese on high until smooth, about 30 seconds.
4 ounces full-fat cream cheese
Beat the sweetened condensed milk into the cream cheese until smooth, about 30 seconds. Set aside.
½ cup sweetened condensed milk
Press about ⅔ of the cake dough mixture into the prepared pan to cover.
Spread the cheesecake mixture evenly over the first dough layer.
Press the remaining ⅓ of the dough into super-flat pieces and lay them over the cheesecake layer to cover as much as possible.
Bake in the preheated oven for 29-31 minutes, or until set in the center and browning along the edges.
Let bars cool in the pan for at least 60 minutes before removing and slicing into 16 squares.